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Butterflies of the Amazon and Andes
 
Variegated Batwing
Achlyodes busirus  CRAMER, 1779
Family - HESPERIIDAE
subfamily - PYRGINAE
Tribe - PYRGINI
 
 introduction | habitats | lifecycle | adult behaviour
 

Achlyodes busirus, male, Boca Manu, Rio Madre de Dios, Peru
 
Introduction
 
The Pyrginae, known colloquially as Flats or Spreadwings, are found in temperate and tropical habitats throughout the world. In the neotropical region there are about 990 described species, of which 581 are placed in the tribe Pyrgini, which includes such well known genera as Urbanus, Astraptes, Bolla, Pyrgus, Erynnis, Sostrata and Mylon. 
Butterflies in the genus Achlyodes are known variously as Batwings, Bat Skippers or Sicklewings. There are 5 known species : minna - found in Bahia, Brazil; mithridates - found on most Caribbean islands and on the mainland from Texas to Peru; pallida - found from Mexico to Bolivia; busirus, which occurs from Costa Rica to Bolivia; and munroei - an exceedingly rare species which was thought to be endemic to Cuba but has recently been discovered in the Bahamas. 
The coloration and pattern varies from species to species, e.g. pallida is light golden brown with dark marbling; mithridates is black with a blue sheen and pale marbling; and busirus is metallic purplish brown with blackish marbling. Achlyodes busirus can in addition be distinguished from related species by the presence of a large yellow patch on the outer half of the underside hindwings.
 
Habitats
 
This is a widespread and common species in much of Central and South America, found at altitudes between sea level and at least 1000m in disturbed habitats including forest edges, glades, clearings, roadsides, riverbanks, orchards and farms.
 
Lifecycle
 
I have no early stage data regarding this species. The following relates to Achlyodes mithridates, and is probably also applicable to busirus : The eggs are laid singly on the upperside of leaves of Citrus, Zanthoxylum and other Rutaceae. The caterpillars vary in colour from yellowish green to bluish grey, and are marked with a dark mid-dorsal line, and a broad band of yellow dashes along the sides. They live solitarily in nests made of silked leaves. The pupa has green wing cases and a yellowish green abdomen.
 
Adult behaviour

 

The butterflies are found throughout the year, but are commonest in the rainy season. During the dry season the species can still be found, but most examples tend to be worn and battered. 

Males are usually seen as singletons amongst mixed mud-puddling aggregations of other Pyrgine skippers such as Anastrus, Antigonus, Ebrietas and Gorgopas. Females are seen less often, and visit flowers for nectar. 

Both sexes rest and feed with the wings outspread, and the forewing apex bent downward.

 

 

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