Butterflies of the Amazon and Andes
Patagonian Grayling
Etcheverrius chiliensis  GUERIN-MENEVILLE, 1830
subfamily - SATYRINAE

Etcheverrius chiliensis, Bariloche, 1100m, Argentina Timothy Boucher
There are 1100 known species of Satyrinae in the neotropical region. About 570 of these are placed in the subtribe Pronophilina - a diverse group of high altitude cloudforest butterflies, all of which are confined to the neotropical region. The vast majority are found only in the Andes, but 4 species are known from the Atlantic cloudforests of Brazil, and there are a further 6 species that are endemic to Guatemala, Costa Rica or Mexico. More oddly there is one genus Calisto that is found exclusively on the Caribbean islands of Cuba and Hispaniola.
Etcheverrius are very similar in appearance and behaviour to the alpine Graylings Oeneis, Chazara and Berberia of the northern hemisphere, which also evolved in cold, arid montane habitats, but are only distantly related. Phylogenetic studies based on DNA sequencing published by Pena, Wahlberg et al in 2006 have shown that Etcheverrius should be placed in the subtribe Pronophilina, a group of high altitude genera endemic to the neotropics.
There are 2 Etcheverrius species - tandilensis which is endemic to Argentina, and chiliensis which is found both in Argentina and Chile.
This species is found on the arid steppes of Patagonia, at altitudes between about 800-1500m.
The lifecycle appears to be unrecorded. The following generalisations are applicable to the subtribe Pronophilina and are probably applicable to Corades : The eggs are round, white or pale greenish white, and laid singly on the foodplants or on surrounding vegetation. The larvae are typically pale brown, marked along the back and sides with narrow dark stripes, and tapering towards each end. The head is large in proportion to the body and has two short forward-pointing horns. The tip of the abdomen is equipped with a pair of caudal prongs which are used to flick the frass away from the feeding area. The larvae of all known Pronophilina feed on Chusquea - a genus of bamboo which grows in thickets, mainly along the courses of streams.
Adult behaviour

To be completed.



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