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Butterflies of the Amazon and Andes
Harmonia Ringlet
Hermeuptychia harmonia  BUTLER, 1867
Family - NYMPHALIDAE
subfamily - SATYRINAE
Tribe - SATYRINI
subtribe - EUPTYCHIINA
Hermeuptychia harmonia, Manu cloudforest, Peru
Introduction
There are about 400 described members of the subtribe Euptychiina, which includes all the neotropical "ringlet" butterflies. Until fairly recently almost all were included in the genus Euptychia, but the revisions by Forster ( 1964 ) and Lamas ( 2004 ) divide this "convenience" genus into a number of smaller genera, including Caeruleuptychia, Cissia, Magneuptychia, Cepheuptychia, Chloreuptychia and Euptychioides.
The "Euptychia" ringlets are notoriously difficult to identify. The ground colour, wingspan, width of the median bands, and size of ocelli vary within each species according to sex, habitat, location, altitude and seasonal climatic conditions. It is appropriate here to quote the words of d'Abrera, who when referring to the hundreds of specimens depicted in his "Butterflies of the Neotropical region" states "I cannot tell with certainty whether the taxa figured are species, races, or infrasubspecific forms".
It has been suggested that the illustration above depicts Hermeuptychia harmonia, and I am listing it as such tentatively, but it is worth noting that the differences between species in this group are so minute that they often escape the attention even of highly experienced taxonomists.  By way of example, in 2009 a "new" species Splendeuptychia ackeryi was separated from other similar species mainly on the basis that it had a slightly "hairier" face !
Hermeuptychia harmonia is found from Colombia to Peru. The insect illustrated above may be a subspecies of harmonia, but could equally be a distinct species.
Habitats
The butterfly depicted on this page was photographed at Manu cloudforest in the eastern Andes of Peru, at an altitudes between 1500m, and has been observed by myself and others at altitudes between about 1400-1900m in the same area.
Lifecycle
Unknown. The lifecycles of related species would seem to indicate that it probably lays it's eggs singly on the foodplant, which is likely to be a member of the Poaceae or Marantaceae. The larvae are likely to be brown, with fine longitudinal lines along the back and sides, and with a pair of caudal prongs.
Adult behaviour

The butterflies are usually encountered in three's and four's in well forested areas, mud-puddling on sand or bare soil in roadside ditches or runnels. They also occur along wide sunlit forest paths, open riverbanks, and other grassy areas.

If disturbed they flutter quickly, but settle within a few metres, usually on grasses or low herbage.

 

 

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