Butterflies of
Britain & Europe
Large Copper
Lycaena dispar batavus
OBERTHUR, 1923
Family - LYCAENIDAE
subfamily -
LYCAENINAE
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Large Copper
Lycaena dispar batavus, male ©
Adrian Hoskins |
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Large Copper
Lycaena dispar batavus, female ©
Adrian Hoskins |
Introduction
The Large Copper is one of Europe's most
beautiful butterflies, the dazzling iridescent copper wings of the
male equalling in brilliance any species found in the tropics. The
female is equally stunning, with a pattern similar to that of the
Small Copper, but having almost double it's wingspan.
The
butterfly was first discovered in 1749 at Dozens Bank in
Lincolnshire, and was originally given the name "Orange Argus of Elloe". The
English subspecies Lycaena dispar dispar
was always considered to be a rare and localised insect,
found only in the Norfolk Broads and on the fens of Cambridgeshire
and Huntingdonshire. Unfortunately for the Large Copper, most of
these habitats, including it's most famous site Whittlesea Mere,
were drained for agriculture - a process that began in the early 17th century.
By the beginning of the 19th century almost all of its breeding sites had been converted to farmland,
and the butterfly was on the verge of extinction. The last known
specimens were captured at Bottisham, Cambridgeshire in 1851.
The
butterfly, in the form of subspecies rutilus,
continued to thrive in parts of Europe however, and is still found
in France, Italy, Germany, Latvia, Poland, southern Finland,
Greece and eastern Turkey. This subspecies was artificially
introduced to a bog in Tipperary, Ireland in 1913, where it
survived for 43 years until the site became overgrown.
In
1915 another race of the butterfly was discovered in the
Netherlands, and given the name Lycaena
dispar batavus. It very closely resembles the original
English race, and was introduced to Woodwalton Fen, a tiny
fragment of fen surrounded by hundreds of square miles of
intensive agriculture, in 1927.
Woodwalton Fen had been specially prepared to receive the Large
Copper, and had been planted with substantial amounts
of the larval foodplant Rumex hydrolapathum.
At first the project was seen as a major success, with over a
thousand adults recorded there in 1928. Unfortunately the site
proved very difficult and costly to manage, and the species
survived there only with considerable assistance from the wardens,
who bred the butterflies in a greenhouse and released batches of
adults every year to boost the failing wild population. It finally
became extinct there in the early 1990's, but may be reintroduced
there or elsewhere in Britain at some stage in the future.
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Large Copper
Lycaena dispar batavus, female ©
Adrian Hoskins |
Habitats
This
species breeds only on wet fenlands and around the boggy margins
of lakes, rivers and canals where the larval foodplant Rumex
hydrolapathum grows in profusion. Suitable habitats no
longer exist in Britain, but the Great Fen Project, which aims to
restore large areas of fenland in eastern England, may eventually
recreate an environment which would justify a further
reintroduction.
In
the Netherlands batavus is found only
at Weerribben and Wieden in Overijjsel, where it breeds around the
margins of lakes formed by peat cutting.
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Large Copper
Lycaena dispar batavus, male ©
Adrian Hoskins |
Lifecycle
In Britain and the Netherlands the butterfly is single-brooded,
emerging in late July and early August, but in warmer parts of the
species range it is double-brooded, emerging in late May and late
August.
The
heavily sculpted eggs are laid singly on leaves of the foodplant,
although sometimes a leaf may be visited several times by one or
more females, resulting in clusters of 3 or 4 eggs, laid in
proximity to the midrib or other prominent ribs. Females
usually avoid ovipositing on plants growing at the water's edge,
choosing instead those that grow in warm sheltered situations
among regenerating vegetation. In Britain and the Netherlands they
are always laid on great water dock R.
hydrolapathum but the subspecies
rutilus ( found elsewhere in Europe ) additionally uses
R. crispus and
R. aquaticus.
The
young caterpillars feed on the underside of the leaves, nibbling
out little grooves but leaving the upper cuticle of the leaf
intact. Larvae of batavus ( and those
of the 2nd brood of rutilus )
hibernate when quite small, attaching themselves to a silken pad
spun on the underside of a dead leaf. They can withstand being
submerged for as long as 3 months during the winter. In the spring
they resume feeding, leaving a distinctive lattice of irregular
shaped holes in the leaf. The fully grown caterpillar is bright
emerald green, smooth in texture, and shaped like a woodlouse.
The
chrysalis is plump, rounded, and buff in colour, with dull greyish
markings along the sides of the abdomen. It is attached by a silk
girdle to a stem of the foodplant.
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Large Copper
Lycaena dispar batavus, female ©
Adrian Hoskins |
Adult behaviour
Males
have a rapid jinking flight just above the herbage, and
periodically settle to bask on thistle or dock leaves. They
spend long periods nectaring at marsh thistles and use them as territorial lookout posts from which they dart
off to investigate passing insects. Females are much more
sedentary in behaviour, spending long periods basking on low
vegetation. I have not observed the courtship but
Thomas states that the pre-nuptial flight is brief, consisting of
the male "flapping his wings over the hen" prior to
copulation.
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