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Text and photographs
protected by Copyright © Adrian Hoskins
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Butterflies of
Britain & Europe
Large Skipper
Ochlodes venata
TURATI, 1905
Family - HESPERIIDAE
subfamily -
HESPERIINAE
introduction
|
habitats |
lifecycle |
adult behaviour
Text and photographs protected by Copyright © Adrian
Hoskins 2007-2008, and must not be reproduced or published in part
or in whole elsewhere in any form without written permission from
Adrian Hoskins. Breach of copyright will be pursued by litigation.
Website designed, produced and owned by
Adrian Hoskins
Large Skipper Ochlodes venata,
male, Glapthorn Cow Pastures, Northamptonshire, England
Introduction
The
Large Skipper, in common with most members of the sub-family Hesperiinae, feeds
in the larval stage on grasses. Another characteristic of the sub-family is the
resting posture as depicted in the illustration, with the forewings held at 45
degrees, and the hindwings held flat. A third characteristic shared with many
other Hesperiines is the presence of a dark streak of androconial ( pheromone
producing ) scales on the forewings of males, as shown above.
This
species is distributed across most of Europe, but absent from northern
Scandinavia, Ireland, much of the southern Iberian peninsula, and most
Mediterranean islands. Beyond Europe it's range extends across temperate Asia as
far as Japan.
There are a colossal number of very similar species found throughout the world.
In Britain however the only similar species is the Silver-spotted Skipper, which
can easily be told apart by the presence on the latter of prominent silvery
spots on the underside hindwings.
Large Skipper Ochlodes venata,
female, Alice Holt Forest, Hampshire, England
Habitats
The butterfly occurs commonly throughout England and
Wales in almost any grassy habitat, but favours damp but sunny sites such as
woodland glades and rides, humid heaths, meadows, and riverside habitats.
Almost all sites are characterised by the presence of bramble
bushes and stands of bracken. Open grassland without bushes or shelter are also used but
populations tend to be much lower.
Large Skipper Ochlodes venata,
male, Stansted Forest
Large Skipper Ochlodes venata,
female, Stansted Forest, West Sussex, England
Lifecycle
The
Large Skipper is single brooded throughout it's range, normally emerging in June
and July, although the flight period in Spain is somewhat longer.
The
pale straw coloured dome-shaped egg is laid singly on the underside of grass blades,
typically on cock's foot or false brome. It hatches after about 10 days.
The
caterpillar feeds on soft lush grasses, favouring cock's foot
Dactylis glomerata,
but will also feed on red fescue
Festuca rubra, false brome
Brachypodium sylvaticum, purple moor grass
Molinea caerulea and other species. It
spends the early stages of it's life within a tube of grass made by joining
together 2 grass blades with strands of silk. It periodically emerges from the
tube to eject it's faeces, which it catapults some distance by flicking with 2
little prongs on it's tail.
It
hibernates within a newly constructed tent of grasses, and resumes feeding in
the spring. When fully grown in early May the larva rests openly on the upperside of grass
blades, but retires to it's grass tube in inclement weather. The mature larva is
green and unmarked, with a purplish black head.
The
chrysalis is dark brown and shiny, with the long detached proboscis case
projecting almost to the tip of the abdomen. It is formed within the larval
shelter, head-upwards, and secured by bristles on the head and cremaster. The
pupal stage lasts about 2 weeks.
Large Skipper Ochlodes venata,
male, Ballard Down, Dorset
Adult behaviour
The Large Skipper has a rapid whirring flight, "skipping" from leaf to leaf. The butterflies settle to bask with wings held in the characteristic position as illustrated, typically on bramble leaves, bracken, or on the flower-heads or blades of cocksfoot and other tall grasses.
Males establish territories in sheltered sunny spots, perching on grass heads, bracken or other herbage. They vigorously defend their territories against other intruding male Large Skippers and against flies and other small rapidly flying insects. During territorial sorties pairs of males buzz frenetically around each other, while flying rapidly in broad circles, soaring to a height of about 4-5 metres above ground level. At this point they separate and the "owner" of the territory returns to it's original perch ( or very nearby ), while the intruding male moves on to set up another territory elsewhere.
I have not observed any form of courtship ritual, and assume that copulation takes place almost immediately after the sexes meet. Copulated pairs can often be found settled on bracken leaves, bramble bushes or coarse grasses in the late morning. When copulated both insects keep their wings closed, and are reluctant to fly, but will do so if disturbed.
At grassland sites the butterflies nectar at bird's foot trefoil, clovers and vetches, but in woodlands they particularly favour bramble blossom and thistles. On heathlands the most common nectar sources are cross-leaved heath and bell heather.
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