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Trip Reports

Butterfly-watching Holidays

Venezuela

October 2000

a group tour organised by Geodyssey, designed and led by Andrew Neild

 

trip report by Adrian Hoskins

 

Venezuela is a highly scenic destination, and has one of the most bio-diverse butterfly faunas in the world. The tour was led by Andrew Neild, the acknowledged world expert on Venezuelan butterflies, so this promised to be an exciting trip. The maximum group size of 8 people however caused me to have strong misgivings about joining the tour, due to the potential difficulties and frustrations of photographing butterflies with so many people in restricted areas of habitat. My fears however proved to be unfounded, as although we often had to queue up to get photographs, everyone remained good natured and we all managed to see and photograph a huge number of species.

Andrew Neild was a very capable, resourceful and amicable tour leader, and was able to show us a huge variety of species, as listed at the end of this report. As well as the butterflies, we saw several attractive Hawkmoths and Silkmoths.

 

While the rest of the group were busy taking photographs, Andrew often wandered about, using a long-poled net to capture canopy dwelling species, some of which were kept cool in papers and subsequently released for photography as dusk approached. He also used tin-foil lures to attract Morphos, and rotting plantains to attract most of the Caligo and Archaeoprepona seen. Several Heraclides, Diaethria, Marpesia and Perisama were attracted to human urine. Many other species were seen mud-puddling, including  about 20 Melete by a stream, about 40 Phoebis around a discarded beer bottle, and another group of about 30 mixed Pieridae that had been attracted to a bright yellow fallen leaf, which they had presumably mistaken for a settled Phoebis.

 

About a dozen new butterfly sub-species, and 4 full species previously unknown to science were captured by Andrew.

 

Birds seen included a variety of hummingbirds, eagles, macaws, nightjars and tanagers. The only mammals we saw alive were squirrels, but we found very fresh pug marks of panther. There were many beautiful dragonflies, damselflies and "helicopters". We came across several snakes : one evening Andrew had a fright when one slithered into his room - fortunately it turned out to be a harmless boa constrictor. Whilst on foot in the savannah of the Gran Sabana we had a very close encounter with a large rattlesnake.

 

The first area we visited was cloudforest at Rio Frio, in the Andes, near the Colombian border. The scenery was very attractive in certain places. The journey to Rio Frio from our hotel at San Cristobal took about 2 hours.  We explored the little used rough road that led alongside the river, and crossed a bridge to reach grassy tracks at higher altitudes. We also explored a track called the Mata Mula trail, and another called La Revancha.

 

The next area visited was some dry forest at Imataca, where we explored along a wide dirt road which cut through the forest. We moved next to the Icabaru road, and later explored forest at Km 80.5 near El Dorado. The following day some of us elected to explore roadside forest on a steep scarp called La Escalera, while others chose to take a short break from butterflies, and walked along a river edge on the Gran Sabana plateau.

 

The group reunited the following day, when we visited Jasper Falls. Finally we took a spectacular flight over the tepuys to Canaima, where we got good views of the world's largest moth Thysania agrippina, resting on a tree trunk. We estimated it's wingspan to be about 11 inches.

 

Before departing for England we each nominated our favourite species of the trip. The candidates included an immaculate male Morpho rhetenor which posed with it's glorious blue wings wide open one afternoon, Hamadryas laodamia, Pterourus zagreus, Dryas julia, Haetera piera, Phoebis philea, Arcas imperialis, Agrias claudina, and such glories as an Archaeoprepona demophoon that sat with it's deep blue-green banded wings half open in full sunshine on a leaf a metre in front of me. One of the most delightful finds was a mating pair of the lovely black, red and white Amarynthis meneria basking with wings spread under a Heliconia leaf.

 

Other candidates included the stunning Nessaea batesii, black, with deep orange and sky-blue wing patches, and a beautiful leaf-green underside; and the gorgeous fresh Morpho hecuba that cruised along a track at eye-level casually flicking its wings in the sunshine. The Red Daggerwing Marpesia petreus with it's wonderfully sculpted bright red-orange wings was another favourite. Another very strong contender was a freshly emerged Caligo atreus which came down to banana bait, and spread open it's deep purple and orange wings before my very privileged eyes.

 

How could I choose a favourite ? Well actually it was very easy - the winner for me was seen at Jasper Falls, a very beautiful place where the river at the base of the falls flows gently, ankle deep, over glistening jasper rock which has been eroded by the river to produce beautiful orange and gold patterns on the surface. In this peaceful idyllic setting, a few minutes ahead of the other group members, I discovered an amazingly beautiful butterfly, a perfect fresh Rhetus periander, it's tailed wings resplendent in the deepest, richest shade of gleaming purple. The butterfly played hide and seek with me for several minutes, dashing from sun-spot to sun-spot in the shady rainforest below the falls. In a state of great excitement I scrambled through the forest trying to relocate this fantastic winged jewel, which taunted me with rare flashes of it's fabulous wings. Then I saw it land for a moment in a tiny spot of bright sunshine, and for two brief but wonderful seconds it spread open it's wings, giving me the greatest privilege any naturalist could imagine.

 

Species recorded in Venezuela in October 2000.

All species listed below were identified either by Andrew Neild in situ, or by Adrian Hoskins, Tony Hoare and Peter Bruce-Jones from circulated photographs, using the d'Abrera volumes, Lewis, Smart, DeVries, Canals, Neild etc. In cases where doubts remained, images were e-mailed to taxonomists specialising in the relevant families, including Jason Hall, Curtis Callaghan, and Andy Warren. Nomenclature follows Neild, and has NOT been updated to follow the more recent revision by Lamas.

Site

Family

subfamily

Genus

species

subspecies

Rio Frio

Hesperiidae

Hesperiinae

Anatrytone

mella

Mata Mula

Hesperiidae

Hesperiinae

Anthoptus

epictetus

epictetus

Km 80.5

Hesperiidae

Hesperiinae

Molo

mango

Mata Mula

Hesperiidae

Hesperiinae

Synapte

malitiosa

pericles

Mata Mula

Hesperiidae

Hesperiinae

Vettius

coryna

catargyra

Rio Frio

Hesperiidae

Hesperiinae

Vettius

coryna

catargyra

Icabaru road

Hesperiidae

Pyrginae

Achlyodes

mithridates

thraso

Rio Frio

Hesperiidae

Pyrginae

Achlyodes

mithridates

thraso

Rio Frio

Hesperiidae

Pyrginae

Antigonus

nearchus

Rio Frio

Hesperiidae

Pyrginae

Arteurotia

tractipennis

contractipennis

Rio Frio

Hesperiidae

Pyrginae

Astraptes

alardus

alardus

Rio Frio

Hesperiidae

Pyrginae

Astraptes

anaphus

anaphus

Imataca

Hesperiidae

Pyrginae

Astraptes

enotrus

Rio Frio

Hesperiidae

Pyrginae

Astraptes

fulgerator

fulgerator

Jasper Falls

Hesperiidae

Pyrginae

Augiades

crinisus

Km 80.5

Hesperiidae

Pyrginae

Autochton

cincta

Imataca

Hesperiidae

Pyrginae

Autochton

cincta

Rio Frio

Hesperiidae

Pyrginae

Autochton

cincta

Rio Frio

Hesperiidae

Pyrginae

Autochton

vectilucis

Rio Frio

Hesperiidae

Pyrginae

Bolla

cupreiceps

Icabaru road

Hesperiidae

Pyrginae

Burca

braco

braco

Mata Mula

Hesperiidae

Pyrginae

Calaenorrhinus

eligius

eligius

Rio Frio

Hesperiidae

Pyrginae

Chioides

catillus

catillus

Jasper Falls

Hesperiidae

Pyrginae

Chiomara

basigutta

basigutta

Icabaru road

Hesperiidae

Pyrginae

Chiomara

basigutta

basigutta

Rio Frio

Hesperiidae

Pyrginae

Cycloglypha

thrasibulus

thrasibulus

Jasper Falls

Hesperiidae

Pyrginae

Discophellus

euribates

euribates

Jasper Falls

Hesperiidae

Pyrginae

Discophellus

porcius

porcius

Rio Frio

Hesperiidae

Pyrginae

Entheus

matho

latifascius

Rio Frio

Hesperiidae

Pyrginae

Gorgopas

chlorocephala

chlorocephala

Rio Frio

Hesperiidae

Pyrginae

Helias

phalaenoides

phalaenoides

Rio Frio

Hesperiidae

Pyrginae

Heliopetes

alana

alana

Rio Frio

Hesperiidae

Pyrginae

Heliopetes

petrus

Icabaru road

Hesperiidae

Pyrginae

Heliopetes

petrus

Imataca

Hesperiidae

Pyrginae

Heliopetes

petrus

Puerto Ordaz

Hesperiidae

Pyrginae

Heliopetes

petrus

La Revancha

Hesperiidae

Pyrginae

Mylon

cajus

cajus

Icabaru road

Hesperiidae

Pyrginae

Mylon

lassia

Rio Frio

Hesperiidae

Pyrginae

Mylon

maimon

Icabaru road

Hesperiidae

Pyrginae

Phanus

vitreus

Imataca

Hesperiidae

Pyrginae

Phanus

vitreus

Rio Frio

Hesperiidae

Pyrginae

Phocides

pigmalion

pigmalion

Rio Frio

Hesperiidae

Pyrginae

Polythrix

hirtius

Rio Frio

Hesperiidae

Pyrginae

Proteides

mercurius

mercurius

Imataca

Hesperiidae

Pyrginae

Pyrgus

oileus

oileus

Km 80.5

Hesperiidae

Pyrginae

Pyrgus

oileus

oileus

La Revancha

Hesperiidae

Pyrginae

Pyrgus

oileus

oileus

Rio Frio

Hesperiidae

Pyrginae

Pyrgus

oileus

oileus

Icabaru road

Hesperiidae

Pyrginae

Pyrgus

oileus

oileus

Mata Mula

Hesperiidae

Pyrginae

Pyrgus

oileus

oileus

Jasper Falls

Hesperiidae

Pyrginae

Pyrgus

oileus

oileus

Jasper Falls

Hesperiidae

Pyrginae

Sophista

aristoteles

aristoteles

Icabaru road

Hesperiidae

Pyrginae

Sostrata

bifasciata

adamas

La Revancha

Hesperiidae

Pyrginae

Theagenes

albiplaga

albiplaga

Rio Frio

Hesperiidae

Pyrginae

Urbanus

chalco

Icabaru road

Hesperiidae

Pyrginae

Urbanus

dorantes

dorantes

Rio Frio

Hesperiidae

Pyrginae

Urbanus

dorantes

dorantes

Rio Frio

Hesperiidae

Pyrginae

Urbanus

proteus

proteus

Mata Mula

Hesperiidae

Pyrginae

Urbanus

proteus

proteus

Icabaru road

Hesperiidae

Pyrginae

Urbanus

proteus

proteus

Rio Frio

Hesperiidae

Pyrginae

Urbanus

simplicius

Rio Frio

Hesperiidae

Pyrginae

Xenophanes

tryxus

Jasper Falls

Hesperiidae

Pyrrhopyginae

Jemadia

gnetus

Icabaru road

Hesperiidae

Pyrrhopyginae

Mysarbia

sejanus

stolli

Rio Frio

Hesperiidae

Pyrrhopyginae

Myscelus

amystis

amystis

Mata Mula

Hesperiidae

Pyrrhopyginae

Pyrrhopyge

decipiens

Rio Frio

Hesperiidae

Pyrrhopyginae

Pyrrhopyge

decipiens

Icabaru road

Hesperiidae

Pyrrhopyginae

Pyrrhopyge

decipiens

La Escalera

Hesperiidae

Pyrrhopyginae

Pyrrhopyge

decipiens

Jasper Falls

Hesperiidae

Pyrrhopyginae

Pyrrhopyge

decipiens

Icabaru road

Lycaenidae

Polyommatinae

Hemiargus

hanno

hanno

Imataca

Lycaenidae

Polyommatinae

Hemiargus

hanno

hanno

Puerto Ordaz

Lycaenidae

Polyommatinae

Hemiargus

hanno

hanno

Icabaru road

Lycaenidae

Polyommatinae

Leptotes

cassius

cassius

Km 80.5

Lycaenidae

Theclinae

Arawacus

aetolus

aetolus

Imataca

Lycaenidae

Theclinae

Arawacus

aetolus

aetolus

Rio Frio

Lycaenidae

Theclinae

Arawacus

aetolus

aetolus

Imataca

Lycaenidae

Theclinae

Arcas

imperialis

Km 80.5

Lycaenidae

Theclinae

Calycopis

cerata

Imataca

Lycaenidae

Theclinae

Calycopis

cerata

Km 80.5

Lycaenidae

Theclinae

Celmia

celmus

Imataca

Lycaenidae

Theclinae

Contrafascia

imma

Imataca

Lycaenidae

Theclinae

Cyanophrys

amyntor

amyntor

Imataca

Lycaenidae

Theclinae

Evenus

coronata

Rio Frio

Lycaenidae

Theclinae

Evenus

gabriela

Mata Mula

Lycaenidae

Theclinae

Lamprospilus

nicetus

Rio Frio

Lycaenidae

Theclinae

Megathecla

cupentus

Mata Mula

Lycaenidae

Theclinae

Micandra

comae

Icabaru road

Lycaenidae

Theclinae

Ministrymon

azia

azia

Rio Frio

Lycaenidae

Theclinae

Paiwarria

episcopalis

Km 80.5

Lycaenidae

Theclinae

Panthiades

bitias

Mata Mula

Lycaenidae

Theclinae

Panthiades

phaleros

Rio Frio

Lycaenidae

Theclinae

Rekoa

meton

Mata Mula

Lycaenidae

Theclinae

Thereus

cithonius

Rio Frio

Nymphalidae

Apaturinae

Doxocopa

cherubina

Rio Frio

Nymphalidae

Apaturinae

Doxocopa

laurentia

chlorotaenia

Rio Frio

Nymphalidae

Apaturinae

Doxocopa

lavinia

Rio Frio

Nymphalidae

Apaturinae

Doxocopa

pavon

pavon

Rio Frio

Nymphalidae

Apaturinae

Doxocopa

zunilda

felderi

Rio Frio

Nymphalidae

Biblidinae

Biblis

hyperia

hyperia

Icabaru road

Nymphalidae

Biblidinae

Callicore

astarte

staudingeri

Rio Frio

Nymphalidae

Biblidinae

Callicore

pitheas

Rio Frio

Nymphalidae

Biblidinae

Catonephele

acontius

acontius

Km 80.5

Nymphalidae

Biblidinae

Catonephele

acontius

acontius

Rio Frio

Nymphalidae

Biblidinae

Catonephele

antinoe

antinoe

Rio Frio

Nymphalidae

Biblidinae

Catonephele

numilia

numilia

Imataca

Nymphalidae

Biblidinae

Catonephele

numilia

numilia

La Revancha

Nymphalidae

Biblidinae

Diaethria

clymena

marchalii

Rio Frio

Nymphalidae

Biblidinae

Diaethria

clymena

clymena

Icabaru road

Nymphalidae

Biblidinae

Diaethria

clymena

clymena

Icabaru road

Nymphalidae

Biblidinae

Diaethria

euclides

metiscus

Rio Frio

Nymphalidae

Biblidinae

Diaethria

gabaza

gabazina

La Revancha

Nymphalidae

Biblidinae

Diaethria

neglecta

merida

La Revancha

Nymphalidae

Biblidinae

Diaethria

neglecta

merida

Mata Mula

Nymphalidae

Biblidinae

Diaethria

neglecta

merida

Rio Frio

Nymphalidae

Biblidinae

Dynamine

anubis

pieridoides

Icabaru road

Nymphalidae

Biblidinae

Dynamine

anubis

anubis

Rio Frio

Nymphalidae

Biblidinae

Dynamine

arene

arene

Rio Frio

Nymphalidae

Biblidinae

Dynamine

postverta

postverta

Km 80.5

Nymphalidae

Biblidinae

Dynamine

racidula

trembathi

Icabaru road

Nymphalidae

Biblidinae

Dynamine

racidula

trembathi

Icabaru road

Nymphalidae

Biblidinae

Eunica

bechina

bechina

Icabaru road

Nymphalidae

Biblidinae

Eunica

concordia

Rio Frio

Nymphalidae

Biblidinae

Eunica

malvina

malvina

Icabaru road

Nymphalidae

Biblidinae

Eunica

mygdonia

mygdonia

Jasper Falls

Nymphalidae

Biblidinae

Eunica

pusilla

Imataca

Nymphalidae

Biblidinae

Eunica

sophonisba

sophonisba

Rio Frio

Nymphalidae

Biblidinae

Haematera

pyrame

thysbe

Rio Frio

Nymphalidae

Biblidinae

Hamadryas

arinome

arienis

Km 80.5

Nymphalidae

Biblidinae

Hamadryas

arinome

arienis

Icabaru road

Nymphalidae

Biblidinae

Hamadryas

februa

februa

Rio Frio

Nymphalidae

Biblidinae

Hamadryas

feronia

farinulenta

Imataca

Nymphalidae

Biblidinae

Hamadryas

feronia

farinulenta

Icabaru road

Nymphalidae

Biblidinae

Hamadryas

velutina

velutina

Rio Frio

Nymphalidae

Biblidinae

Marpesia

berania

berania

Icabaru road

Nymphalidae

Biblidinae

Marpesia

berania

berania

Icabaru road

Nymphalidae

Biblidinae

Marpesia

chiron

Imataca

Nymphalidae

Biblidinae

Marpesia

chiron

Jasper Falls

Nymphalidae

Biblidinae

Marpesia

chiron

Rio Frio

Nymphalidae

Biblidinae

Marpesia

chiron

Rio Frio

Nymphalidae

Biblidinae

Marpesia

corinna

Icabaru road

Nymphalidae

Biblidinae

Marpesia

furcula

oechalia

La Revancha

Nymphalidae

Biblidinae

Marpesia

marcella

marcella

Jasper Falls

Nymphalidae

Biblidinae

Marpesia

petreus

petreus

Rio Frio

Nymphalidae

Biblidinae

Marpesia

petreus

petreus

Icabaru road

Nymphalidae

Biblidinae

Marpesia

petreus

petreus

Imataca

Nymphalidae

Biblidinae

Marpesia

petreus

petreus

Icabaru road

Nymphalidae

Biblidinae

Marpesia

themistocles

La Revancha

Nymphalidae

Biblidinae

Marpesia

zerynthia

dentigera

Icabaru road

Nymphalidae

Biblidinae

Mestra

dorcas

hersilia

Imataca

Nymphalidae

Biblidinae

Mestra

dorcas

hersilia

Rio Frio

Nymphalidae

Biblidinae

Mestra

dorcas

hersilia

Rio Frio

Nymphalidae

Biblidinae

Myscelia

cyaniris

albescens

Imataca

Nymphalidae

Biblidinae

Myscelia

cyaniris

albescens

Km 80.5

Nymphalidae

Biblidinae

Nessaea

batesii

magniplaga

Km 80.5

Nymphalidae

Biblidinae

Nessaea

obrinus

lesoudieri

Rio Frio

Nymphalidae

Biblidinae

Nica

flavilla

canthara

La Revancha

Nymphalidae

Biblidinae

Orophila

diotima

cecidas

Imataca

Nymphalidae

Biblidinae

Peria

lamis

La Revancha

Nymphalidae

Biblidinae

Perisama

bomplandii

parabomplandii

La Revancha

Nymphalidae

Biblidinae

Perisama

dorbignyi

dorbignyi

La Revancha

Nymphalidae

Biblidinae

Perisama

humboldtii

tringa

La Revancha

Nymphalidae

Biblidinae

Perisama

oppelii

angustifasciata

La Revancha

Nymphalidae

Biblidinae

Perisama

yeba

guerini

Km 80.5

Nymphalidae

Biblidinae

Pyrrhogyra

amphiro

juani

Imataca

Nymphalidae

Biblidinae

Pyrrhogyra

neaerea

neaerea

Rio Frio

Nymphalidae

Biblidinae

Pyrrhogyra

neaerea

kheili

Jasper Falls

Nymphalidae

Biblidinae

Temenis

laothoe

laothoe

Mata Mula

Nymphalidae

Biblidinae

Temenis

laothoe

colombiana

Km 80.5

Nymphalidae

Biblidinae

Temenis

laothoe

laothoe

La Revancha

Nymphalidae

Biblidinae

Temenis

laothoe

colombiana

Icabaru road

Nymphalidae

Biblidinae

Temenis

laothoe

laothoe

Rio Frio

Nymphalidae

Biblidinae

Temenis

laothoe

colombiana

Jasper Falls

Nymphalidae

Charaxinae

Agrias

amydon

ssp. nov 1

Jasper Falls

Nymphalidae

Charaxinae

Agrias

claudina

sardanapalus

Jasper Falls

Nymphalidae

Charaxinae

Archaeoprepona

amphimachus

amphimachus

La Revancha

Nymphalidae

Charaxinae

Archaeoprepona

amphimachus

amphimachus

Km 80.5

Nymphalidae

Charaxinae

Archaeoprepona

amphimachus

amphimachus

Rio Frio

Nymphalidae

Charaxinae

Archaeoprepona

amphimachus

amphimachus

Icabaru road

Nymphalidae

Charaxinae

Archaeoprepona

amphimachus

amphimachus

Rio Frio

Nymphalidae

Charaxinae

Archaeoprepona

demophon

demophon

Km 80.5

Nymphalidae

Charaxinae

Archaeoprepona

demophon

demophon

Jasper Falls

Nymphalidae

Charaxinae

Archaeoprepona

demophon

demophon

Icabaru road

Nymphalidae

Charaxinae

Archaeoprepona

demophon

demophon

Icabaru road

Nymphalidae

Charaxinae

Archaeoprepona

demophoon

andicola

Rio Frio

Nymphalidae

Charaxinae

Archaeoprepona

demophoon

andicola

Jasper Falls

Nymphalidae

Charaxinae

Archaeoprepona

licomedes

licomedes

Icabaru road

Nymphalidae

Charaxinae

Archaeoprepona

licomedes

licomedes

Rio Frio

Nymphalidae

Charaxinae

Consul

fabius

bogotanus

Km 80.5

Nymphalidae

Charaxinae

Consul

fabius

bogotanus

Rio Frio

Nymphalidae

Charaxinae

Fountainea

ryphea

ryphea

Icabaru road

Nymphalidae

Charaxinae

Fountainea

ryphea

ryphea

Icabaru road

Nymphalidae

Charaxinae

Hypna

clytemnestra

Imataca

Nymphalidae

Charaxinae

Hypna

clytemnestra

Rio Frio

Nymphalidae

Charaxinae

Hypna

clytemnestra

Icabaru road

Nymphalidae

Charaxinae

Memphis

glauce

glauce

El Palmar

Nymphalidae

Charaxinae

Memphis

laertes

La Escalera

Nymphalidae

Charaxinae

Memphis

montesino

Icabaru road

Nymphalidae

Charaxinae

Memphis

montesino

Icabaru road

Nymphalidae

Charaxinae

Memphis

moruus

moruus

Icabaru road

Nymphalidae

Charaxinae

Memphis

oenomais

Rio Frio

Nymphalidae

Charaxinae

Memphis

philumena

chaeronia

Km 80.5

Nymphalidae

Charaxinae

Memphis

polycarmes

Rio Frio

Nymphalidae

Charaxinae

Memphis

xenocles

xenocles

Icabaru road

Nymphalidae

Charaxinae

Polygrapha

xenocrates

punctimarginale

Km 80.5

Nymphalidae

Charaxinae

Prepona

laertes

louisa

Rio Frio

Nymphalidae

Charaxinae

Prepona

laertes

louisa

Imataca

Nymphalidae

Charaxinae

Prepona

laertes

louisa

Icabaru road

Nymphalidae

Charaxinae

Prepona

laertes

louisa

Icabaru road

Nymphalidae

Charaxinae

Prepona

laertes

louisa

Km 80.5

Nymphalidae

Charaxinae

Prepona

pheridamas

Rio Frio

Nymphalidae

Charaxinae

Prepona

pylene

eugenes

Icabaru road

Nymphalidae

Charaxinae

Siderone

galanthis

galanthis

Km 80.5

Nymphalidae

Charaxinae

Zaretis

itys

itys

Imataca

Nymphalidae

Danainae

Danaus

plexippus

plexippus

Rio Frio

Nymphalidae

Danainae

Danaus

plexippus

plexippus

Icabaru road

Nymphalidae

Danainae

Lycorea

halia

atergatis

Rio Frio

Nymphalidae

Danainae

Lycorea

halia

atergatis

La Revancha

Nymphalidae

Danainae

Lycorea

ilione

lamirus

Rio Frio

Nymphalidae

Danainae

Lycorea

ilione

lamirus

La Revancha

Nymphalidae

Heliconiinae

Abananote

hylonome

basilia

Mata Mula

Nymphalidae

Heliconiinae

Abananote

hylonome

basilia

Mata Mula

Nymphalidae

Heliconiinae

Actinote

anteas

anteas

Rio Frio

Nymphalidae

Heliconiinae

Actinote

pellenea

hahneli

Icabaru road

Nymphalidae

Heliconiinae

Actinote

thalia

thalia

Jasper Falls

Nymphalidae

Heliconiinae

Agraulis

vanillae

vanillae

Icabaru road

Nymphalidae

Heliconiinae

Agraulis

vanillae

vanillae

La Revancha

Nymphalidae

Heliconiinae

Altinote

dicaeus

dicaeus

Mata Mula

Nymphalidae

Heliconiinae

Altinote

dicaeus

dicaeus

Icabaru road

Nymphalidae

Heliconiinae

Dione

juno

juno

Icabaru road

Nymphalidae

Heliconiinae

Dryas

iulia

alcionea

Rio Frio

Nymphalidae

Heliconiinae

Dryas

iulia

alcionea

Jasper Falls

Nymphalidae

Heliconiinae

Dryas

iulia

alcionea

Mata Mula

Nymphalidae

Heliconiinae

Dryas

iulia

alcionea

Km 80.5

Nymphalidae

Heliconiinae

Dryas

iulia

alcionea

Rio Frio

Nymphalidae

Heliconiinae

Eueides

aliphera

gracilis

Km 80.5

Nymphalidae

Heliconiinae

Eueides

aliphera

gracilis

Rio Frio

Nymphalidae

Heliconiinae

Eueides

isabella

dynastes

Icabaru road

Nymphalidae

Heliconiinae

Eueides

lybia

orinocensis

La Revancha

Nymphalidae

Heliconiinae

Eueides

procula

Rio Frio

Nymphalidae

Heliconiinae

Eueides

tales

franciscus

Rio Frio

Nymphalidae

Heliconiinae

Eueides

vibilia

Icabaru road

Nymphalidae

Heliconiinae

Eueides

vibilia

La Revancha

Nymphalidae

Heliconiinae

Heliconius

antiochus

araneides

Rio Frio

Nymphalidae

Heliconiinae

Heliconius

antiochus

araneides

Rio Frio

Nymphalidae

Heliconiinae

Heliconius

charithonia

charithonia

Mata Mula

Nymphalidae

Heliconiinae

Heliconius

clysonymus

clysonymus

La Revancha

Nymphalidae

Heliconiinae

Heliconius

cydno

Km 80.5

Nymphalidae

Heliconiinae

Heliconius

egeria

keithbrowni

Icabaru road

Nymphalidae

Heliconiinae

Heliconius

elevatus

roraima

Imataca

Nymphalidae

Heliconiinae

Heliconius

erato

dignus

Icabaru road

Nymphalidae

Heliconiinae

Heliconius

erato

dignus

Km 80.5

Nymphalidae

Heliconiinae

Heliconius

erato

dignus

Jasper Falls

Nymphalidae

Heliconiinae

Heliconius

erato

dignus

Rio Frio

Nymphalidae

Heliconiinae

Heliconius

erato

dignus

Rio Frio

Nymphalidae

Heliconiinae

Heliconius

hecale

anderina

Rio Frio

Nymphalidae

Heliconiinae

Heliconius

melpomene

bellula

Jasper Falls

Nymphalidae

Heliconiinae

Heliconius

melpomene

melpomene

Icabaru road

Nymphalidae

Heliconiinae

Heliconius

melpomene

melpomene

Imataca

Nymphalidae

Heliconiinae

Heliconius

melpomene

melpomene

Rio Frio

Nymphalidae

Heliconiinae

Heliconius

numata

messene

Km 80.5

Nymphalidae

Heliconiinae

Heliconius

numata

numata

Rio Frio

Nymphalidae

Heliconiinae

Heliconius

sara

magdalena

Icabaru road

Nymphalidae

Heliconiinae

Heliconius

sara

sara

Km 80.5

Nymphalidae

Heliconiinae

Heliconius

sara

sara

Mata Mula

Nymphalidae

Heliconiinae

Heliconius

sara

magdalena

Jasper Falls

Nymphalidae

Heliconiinae

Heliconius

sara

sara

Km 80.5

Nymphalidae

Heliconiinae

Heliconius

wallacei

flavescens

Icabaru road

Nymphalidae

Heliconiinae

Heliconius

wallacei

flavescens

Imataca

Nymphalidae

Heliconiinae

Heliconius

wallacei

wallacei

Icabaru road

Nymphalidae

Heliconiinae

Heliconius

xanthocles

cleoxanthe

Rio Frio

Nymphalidae

Heliconiinae

Laparus

doris

dives

Caracas

Nymphalidae

Heliconiinae

Laparus

doris

dives

Km 80.5

Nymphalidae

Heliconiinae

Laparus

doris

dives

Km 80.5

Nymphalidae

Heliconiinae

Neruda

aoede

ayacuchensis

Rio Frio

Nymphalidae

Heliconiinae

Philaethria

dido

dido

Icabaru road

Nymphalidae

Heliconiinae

Philaethria

dido

dido

Imataca

Nymphalidae

Heliconiinae

Philaethria

dido

dido

Mata Mula

Nymphalidae

Heliconiinae

Podotricha

judith

judith

Icabaru road

Nymphalidae

Ithomiinae

Callithomia

lenea

lenea

La Escalera

Nymphalidae

Ithomiinae

Callithomia

lenea

lenea

Icabaru road

Nymphalidae

Ithomiinae

Ceratinia

neso

nisea

Rio Frio

Nymphalidae

Ithomiinae

Ceratinia

neso

hamlini

Rio Frio

Nymphalidae

Ithomiinae

Ceratinia

tutia

La Escalera

Nymphalidae

Ithomiinae

Dircenna

adina

adina

La Escalera

Nymphalidae

Ithomiinae

Dircenna

dero

euchytma

La Revancha

Nymphalidae

Ithomiinae

Hypoleria

ocalea

Icabaru road

Nymphalidae

Ithomiinae

Hypothyris

euclea

euclea

Rio Frio

Nymphalidae

Ithomiinae

Hypothyris

euclea

euclea

La Escalera

Nymphalidae

Ithomiinae

Hypothyris

ninonia

mysotis

Icabaru road

Nymphalidae

Ithomiinae

Hypothyris

ninonia

mysotis

Rio Frio

Nymphalidae

Ithomiinae

Hypothyris

ninonia

mysotis

Rio Frio

Nymphalidae

Ithomiinae

Ithomia

iphianassa

iphianassa

La Revancha

Nymphalidae

Ithomiinae

Ithomia

terra

terra

Icabaru road

Nymphalidae

Ithomiinae

Mechanitis

lysimnia

lysimnia

Km 80.5

Nymphalidae

Ithomiinae

Mechanitis

mazaeus

beebei

Rio Frio

Nymphalidae

Ithomiinae

Mechanitis

mazaeus

fallax

Km 80.5

Nymphalidae

Ithomiinae

Mechanitis

polymnia

polymnia

Icabaru road

Nymphalidae

Ithomiinae

Mechanitis

polymnia

polymnia

Rio Frio

Nymphalidae

Ithomiinae

Mechanitis

polymnia

veritabilis

Icabaru road

Nymphalidae

Ithomiinae

Melinaea

ludovica

ludovica

Imataca

Nymphalidae

Ithomiinae

Melinaea

mneme

mauensis

Rio Frio

Nymphalidae

Ithomiinae

Melinaea

mneme

mauensis

La Escalera

Nymphalidae

Ithomiinae

Napeogenes

rhezia

adulta

Icabaru road

Nymphalidae

Ithomiinae

Oleria

astrea

thiemei

Mata Mula

Nymphalidae

Ithomiinae

Oleria

makrena

Mata Mula

Nymphalidae

Ithomiinae

Oleria

makrena

La Escalera

Nymphalidae

Ithomiinae

Oleria

santinesa

Mata Mula

Nymphalidae

Ithomiinae

Pteronymia

alida

alida

La Escalera

Nymphalidae

Ithomiinae

Pteronymia

artena

beebei

Rio Frio

Nymphalidae

Ithomiinae

Pteronymia

latilla

nigricans

La Escalera

Nymphalidae

Ithomiinae

Pteronymia

oneida

asopo

Icabaru road

Nymphalidae

Ithomiinae

Pteronymia

primula

primula

Icabaru road

Nymphalidae

Ithomiinae

Pteronymia

sao

sao

Rio Frio

Nymphalidae

Ithomiinae

Pteronymia

sp.nov 1 Neild

La Escalera

Nymphalidae

Ithomiinae

Scada

reckia

philemon

Icabaru road

Nymphalidae

Ithomiinae

Thyridia

psidii

aedesia

La Revancha

Nymphalidae

Ithomiinae

Thyridia

psidii

aedesia

Rio Frio

Nymphalidae

Ithomiinae

Tithorea

harmonia

furia

Rio Frio

Nymphalidae

Ithomiinae

Tithorea

tarricina

hecalesina

Rio Frio

Nymphalidae

Limenitidinae

Adelpha

capucinus

capucinus

Imataca

Nymphalidae

Limenitidinae

Adelpha

cocala

orellanae

Rio Frio

Nymphalidae

Limenitidinae

Adelpha

cytherea

nahua

Km 80.5

Nymphalidae

Limenitidinae

Adelpha

cytherea

nahua

Icabaru road

Nymphalidae

Limenitidinae

Adelpha

cytherea

nahua

La Revancha

Nymphalidae

Limenitidinae

Adelpha

delinita

delinata

Rio Frio

Nymphalidae

Limenitidinae

Adelpha

iphiclus

iphiclus

Jasper Falls

Nymphalidae

Limenitidinae

Adelpha

iphiclus

iphiclus

Icabaru road

Nymphalidae

Limenitidinae

Adelpha

iphiclus

iphiclus

La Escalera

Nymphalidae

Limenitidinae

Adelpha

irmina

irmina

Mata Mula

Nymphalidae

Limenitidinae

Adelpha

justina

justinella

Rio Frio

Nymphalidae

Limenitidinae

Adelpha

lycorias

lara

Rio Frio

Nymphalidae

Limenitidinae

Adelpha

malea

malea

Km 80.5

Nymphalidae

Limenitidinae

Adelpha

melona

leucoma

Jasper Falls

Nymphalidae

Limenitidinae

Adelpha

melona

leucoma

Jasper Falls

Nymphalidae

Limenitidinae

Adelpha

mesentina

Mata Mula

Nymphalidae

Limenitidinae

Adelpha

seriphia

La Escalera

Nymphalidae

Morphinae

Antirrhea

philoctetes

ulei

Km 80.5

Nymphalidae

Morphinae

Bia

actorion

actorion

Rio Frio

Nymphalidae

Morphinae

Caligo

atreus

ajax

La Revancha

Nymphalidae

Morphinae

Caligo

eurilochus

delectans

Km 80.5

Nymphalidae

Morphinae

Caligo

eurilochus

delectans

Icabaru road

Nymphalidae

Morphinae

Caligo

eurilochus

delectans

Icabaru road

Nymphalidae

Morphinae

Caligo

idomeneus

idomenides

Rio Frio

Nymphalidae

Morphinae

Caligo

idomeneus

idomenides

Icabaru road

Nymphalidae

Morphinae

Caligo

illioneus

illioneus

Rio Frio

Nymphalidae

Morphinae

Caligo

illioneus

illioneus

Rio Frio

Nymphalidae

Morphinae

Caligo

oileus

oileus

Km 80.5

Nymphalidae

Morphinae

Caligo

suzanna

Icabaru road

Nymphalidae

Morphinae

Catoblepia

berecynthia

halli

Icabaru road

Nymphalidae

Morphinae

Catoblepia

soranus

Jasper Falls

Nymphalidae

Morphinae

Eryphanis

automedon

automedon

Rio Frio

Nymphalidae

Morphinae

Morpho

achilles

guaraunos

Imataca

Nymphalidae

Morphinae

Morpho

achilles

guaraunos

Jasper Falls

Nymphalidae

Morphinae

Morpho

achilles

guaraunos

Icabaru road

Nymphalidae

Morphinae

Morpho

achilles

guaraunos

Icabaru road

Nymphalidae

Morphinae

Morpho

cisseis

cisseis

Jasper Falls

Nymphalidae

Morphinae

Morpho

deidamia

guaraura

Icabaru road

Nymphalidae

Morphinae

Morpho

deidamia

guaraura

Rio Frio

Nymphalidae

Morphinae

Morpho

deidamia

guaraura

Imataca

Nymphalidae

Morphinae

Morpho

deidamia

guaraura

Icabaru road

Nymphalidae

Morphinae

Morpho

hecuba

polyidos

Km 80.5

Nymphalidae

Morphinae

Morpho

hecuba

polyidos

Rio Frio

Nymphalidae

Morphinae

Morpho

helenor

tucupita

Imataca

Nymphalidae

Morphinae

Morpho

helenor

tucupita

Icabaru road

Nymphalidae

Morphinae

Morpho

helenor

tucupita

Jasper Falls

Nymphalidae

Morphinae

Morpho

helenor

peleides

Rio Frio

Nymphalidae

Morphinae

Morpho

marcus

intermedia

Icabaru road